Des Moinеs (USA), 28 May 1942

Foreign member

Department of Chemical and Biological Sciences


Membership of SASA
Foreign member since 30 October 2003

Field of study

BSc (1964); MSc (1968), University of Pennsylvania

University of California San Francisco, professor (1984‒); Institute for Neurodegenerative Diseases, director (1999‒)

Membership of specialist associations
American Society of Biochemistry and Molecular Biology; American Society for Clinical Investigation, emeritus; American Chemical Society, American Association for the Advancement of Science; American Academy of Neurology, fellow; American Society for Neurochemistry, honorary member; American Society for Microbiology, fellow; International Society for Neurochemistry; American Society for Cell Biology; California Medical Association; San Francisco Neurological Society; Society for Neuroscience; American Neurological Association, honorary member; American Society of Virology; American Society of Human Genetics; Association of American Physicians; Association of California Neurologists; American Association of Anatomists, honorary member; American Society of Neuroradiology, honorary member

Awards and recognitions
National Medal of Science, USA (2009);
Nobel Prize in medicine (1997);
Wolf-Prize (1996);
Albert Lasker Award for Basic Medical Research (1994);
the Gairdner Foundation International Award (1993);
Potamkin Prize from the American Academy of Neurology (1991)